Nathan Fillion

Nathan Fillion (born March 27, 1971) is a Canadian-American actor. He played the leading roles of Captain Malcolm "Mal" Reynolds on Firefly and its film continuation Serenity and Richard Castle on Castle. As of 2018, he stars as John Nolan on The Rookie and is an executive producer on the show as well as its spin-off series, The Rookie: Feds.

Fillion has acted in traditionally distributed films like Slither and Trucker, Internet-distributed films like Dr. Horrible's Sing-Along Blog, television soap operas, sitcoms, and theatre. His voice is featured in animation and video games, such as the Bungiegames Halo 3, Halo 3: ODST, Halo: Reach, Destiny, and Destiny 2, along with the 343 Industries game Halo 5: Guardians and the television series M.O.D.O.K. (2021).

Fillion first gained recognition for his work on One Life to Live in the contract role of Joey Buchanan, for which he was nominated for the Daytime Emmy Award for Outstanding Younger Actor in a Drama Series, as well as for his supporting role as Johnny Donnelly in the sitcom Two Guys and a Girl.

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Nathan Fillion has built a filmography of 20 credits over a career spanning from 1997 to 2025.

Nathan Fillion's most notable role to date came in Guardians of the Galaxy (2014), where he played Monstrous Inmate. He also appeared in Saving Private Ryan (1998) as Minnesota Ryan. Most recently appeared in Monsters University (2013) as Johnny (voice).

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